①College of Biology and Environment, Nanjing Forestry University, Nanjing 210037; ②College of Life and Environment Sciences, Huangshan University, Huangshan 245041,③Guangdong Entomological Institute, Guangzhou, 510260;④Guangdong Public Laboratory of Wild Animal Conservation and Utilization, Guangzhou 510260,①College of Biology and Environment, Nanjing Forestry University, Nanjing 210037,③Guangdong Entomological Institute, Guangzhou, 510260;④Guangdong Public Laboratory of Wild Animal Conservation and Utilization, Guangzhou 510260,②College of Life and Environment Sciences, Huangshan University, Huangshan 245041