Institute of Hydorbiology, Jinan University, Key Laboratory of Aquatic Eutrophication and Control of Harmful Algal Blooms of Guangdong Higher Education Institutes ,Guangzhou 510632,China,Institute of Hydorbiology, Jinan University, Key Laboratory of Aquatic Eutrophication and Control of Harmful Algal Blooms of Guangdong Higher Education Institutes ,Guangzhou 510632,China,Institute of Hydorbiology, Jinan University, Key Laboratory of Aquatic Eutrophication and Control of Harmful Algal Blooms of Guangdong Higher Education Institutes ,Guangzhou 510632,China,Institute of Hydorbiology, Jinan University, Key Laboratory of Aquatic Eutrophication and Control of Harmful Algal Blooms of Guangdong Higher Education Institutes ,Guangzhou 510632,China,Institute of Hydorbiology, Jinan University, Key Laboratory of Aquatic Eutrophication and Control of Harmful Algal Blooms of Guangdong Higher Education Institutes ,Guangzhou 510632,China,Institute of Hydorbiology, Jinan University, Key Laboratory of Aquatic Eutrophication and Control of Harmful Algal Blooms of Guangdong Higher Education Institutes ,Guangzhou 510632,China