①College of Life and Environment Sciences, Huangshan University, Huangshan 245041; ②Bioresource institute, Huangshan University, Huangshan 245041; ③Graduate School of Nanjing Forestry University, Nanjing 210037,②Bioresource institute, Huangshan University, Huangshan 245041,①College of Life and Environment Sciences, Huangshan University, Huangshan 245041; ②Bioresource institute, Huangshan University, Huangshan 245041