Beijing Zoo,Administration of Fanjingshan National Nature Reserve, Tongren, Guizhou,San Diego Zoo Institute for Conservation Research, Escondido, CA 92027 USA,German Primate Center,Administration of Fanjingshan National Nature Reserve, Tongren, Guizhou, China,Administration of Fanjingshan National Nature Reserve, Tongren, Guizhou, China,Beijing Zoo,Administration of Fanjingshan National Nature Reserve, Tongren, Guizhou, China